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FAMILY MYCETOPHILIDAE
Paul L.Th. Beuk Version date: 15-11-2023 15:30

Leia bimaculata (Meigen, 1804)
Diagnosis. Minute to rather large (2.0-15.0 mm) fungus gnats, body varying from pale to dark. Head small; two or three ocelli, lateral ocelli sometimes located next to compound eyes; antennae short to long, segments sometimes close together resembling a single third segment. Thorax hump-backed; wings clear or with more or less extensive markings consisting of smudges, spots or fasciae, cross-vein bm-cu (if present) very close to base of wing, radius with two or three branches, second branch sometimes merging with R1 to form closed cell in anterior half of wing; legs short to long, tarsal segments sometimes slightly swollen in females. Abdomen short to long, sometimes swollen or compressed laterally or dorsoventrally.

Biology. Most reared species were associated with a wide variety of fungi, both Ascomycetina and Basidiomycetina, most others were associated with dead wood. The species associated with dead wood mostly live on the dead wood (or behind bark) rather than in the dead wood. It is suggested that most of the species not directly associated with fungi live on mycelia in the substrates. Adults are most often found in dark humid places, often near fallen trees, dead wood, banks, cliffs, etc.

General references. Chandler (1980 [Ectrepesthoneura], 1981 [Epicypta]), Gagné (1975 [Phronia], 1981b [Trichonta]), Hackman et al. (1988 [catalogue]), Hutson (1979 [Syntemna]), Hutson et al. (1980 [keys Gnoristinae, Leiinae, Mycomyinae, Sciophilinae]), Laštovka (1972 [Mycetophila ruficollis-group]), Väisänen (1982a [Neoempheria], 1984 [Mycomya]), Zaitsev (1983a [Monoclona], 1983b [Allodia s.str.], 1986 [Dynatosoma], 1989a [Zygomyia], 1989b [Anatella]).

References to the local fauna. Barendrecht (1938), Bechev (1997), Beuk (1996h, 1997), Beuk et al. (1994, 1995), Burger et al. (1984), Hackman et al. (1988), De Meijere (1935, 1939a, 1939b), Meuffels (1970), Plassmann (1970), Väisänen (1982b, 1984), Van der Wulp (1874, 1877), Van der Wulp & De Meijere (1898).

How to quote this page: Beuk, P.L.Th., 2023. Family Mycetophilidae. In: Beuk, P.L.Th. (Ed.): Checklist of the Diptera of the Netherlands, https://diptera-info.nl/news.php?fam=Mycetophilidae (date accessed: 22/04/2024).

NL: 255, 2I, 4? B: 161 + 1? D: 251, 1I, 3? UK: 476 World: 4000
 
MYCETOPHILIDAE
 
GNORISTINAE
                 
  Apolephthisa Grzegorzek, 1885
  subincana (Curtis, 1837) N; will be formally added by Beuk & Chandler (in prep.) 
                 
  Boletina Staeger, 1840
  basalis (Meigen, 1818)
  dispecta Dziedzicki, 1885 Note 1
  dubia (Meigen, 1804)
  gripha Dziedzicki, 1885
  griphoides Edwards, 1925
  nitida Grzegorzek, 1885 N; will be formally added by Beuk & Chandler (in prep.) 
  plana (Walker, 1856)
  sciarina Staeger, 1840
  silvatica Dziedzicki, 1885
  trivittata (Meigen, 1818)
                 
  Coelosia Winnertz, 1863
  flava (Staeger, 1840) N; will be formally added by Beuk & Chandler (in prep.) 
  fusca Bezzi, 1892 N; occurrence following Fauna Europaea but it was already listed as silvatica by Beuk (2002e
  = silvatica Landrock, 1918 Note 2
  tenella (Zetterstedt, 1852)
                 
  Synapha Meigen, 1818
  = Empalia Winnertz, 1863
  fasciata Meigen, 1818 N; added by Beuk (1997: xxii
  vitripennis (Meigen, 1818)
                 
  Syntemna Winnertz, 1863
  stylata Hutson, 1979 N; will be formally added by Beuk & Chandler (in prep.) 
 
LEIINAE
                 
  Clastobasis Skuse, 1890
  alternans (Winnertz, 1863) N; will be formally added by Beuk & Chandler (in prep.) 
                 
  Docosia Winnertz, 1863
  fuscipes (von Roser, 1840) N; will be formally added by Beuk & Chandler (in prep.) 
  gilvipes (Walker, 1856)
  pallipes Edwards, 1941 N; will be formally added by Beuk & Chandler (in prep.) 
  sciarina (Meigen, 1830)
  = valida Winnertz, 1863
                 
  Ectrepesthoneura Enderlein, 1911
  hirta (Winnertz, 1846)
                 
  Leia Meigen, 1818
  = Glaphyroptera Winnertz, 1863: preocc.
  = Neoglaphyroptera Osten Sacken, 1878
  arsona Hutson, 1978 I; added by Burger et al. (1984: 145
  bimaculata (Meigen, 1804)
  = fasciola Meigen, 1818
  = stigmatella Van der Wulp, 1859
  fascipennis Meigen, 1818
  longiseta Barendrecht, 1938
  picta Meigen, 1830
  subfasciata (Meigen, 1818)
  winthemi Lehmann, 1822
                 
  Megophthalmidia Dziedzicki, 1889
  crassicornis (Curtis, 1837) Note 3
  = ferruginea (Van der Wulp, 1859)
  = zugmayeriae Dziedzicki, 1889
                 
  Rondaniella Johannsen, 1909
  dimidiata (Meigen, 1804)
  = variegata (Winnertz, 1863)
                 
  Tetragoneura Winnertz, 1846
  sylvatica Curtis, 1837
  = distincta Winnertz, 1846
 
MYCETOPHILINAE
 
  Exechiini
                 
  Allodia Winnertz, 1863
  anglofennica Edwards, 1921
  lugens (Wiedemann, 1817)
  lundstroemi Edwards, 1921
  ornaticollis (Meigen, 1818)
  = nigricollis (Zetterstedt, 1852)
  = longicornis (Van der Wulp, 1859)
  truncata Edwards, 1921
                 
  Allodiopsis Tuomikoski, 1966
  domestica (Meigen, 1830)
  rustica (Edwards, 1941) N; will be formally added by Beuk & Chandler (in prep.) 
                 
  Anatella Winnertz, 1863
  ciliata Winnertz, 1863
  flavomaculata Lundström, 1909 N; occurrence according to Fauna Europaea  Note 4
  longisetosa Dziedzicki, 1923 N; added by Beuk (2021b: 19
  schmitzi Landrock, 1925
  unguigera Edwards, 1921
                 
  Brachycampta Winnertz, 1863
  alternans (Zetterstedt, 1838)
  barbata (Lundström, 1909) N; will be formally added by Beuk & Chandler (in prep.) 
  grata (Meigen, 1830)
  pistillata (Lundström, 1911) N; will be formally added by Beuk & Chandler (in prep.) 
                 
  Brevicornu Marshall, 1869
  arcticoides Caspers, 1985 N; occurrence according to Fauna Europaea 
  auriculatum (Edwards, 1925) N; will be formally added by Beuk & Chandler (in prep.) 
  fissicauda (Lundström, 1911)
  fuscipenne (Staeger, 1840) N; will be formally added by Beuk & Chandler (in prep.)  Note 5
  griseicolle (Staeger, 1840)
  = caudata (Winnertz, 1863)
  intermedium (Santos Abreu, 1920) N; occurrence according to Fauna Europaea 
  nigrofuscum (Lundström, 1909) N; occurrence according to Fauna Europaea 
  ruficorne (Meigen, 1838)
  = cinereum (Lundström, 1911)
  sericoma (Meigen, 1830)
  = amoena (Winnertz, 1863)
                 
  Cordyla Meigen, 1803
  = Polyxena Meigen, 1800: suppr.
  bicornuta (Landrock, 1926) [T] 
  brevicornis (Staeger, 1840)
  crassicornis Meigen, 1818
  = cinerea (Zetterstedt, 1852)
  fasciata Meigen, 1830
  fissa Edwards, 1925
  flaviceps (Staeger, 1840)
  fusca Meigen, 1804
  = atra (Macquart, 1834)
  murina Winnertz, 1863
  nitidula Edwards, 1925 Note 6
  = bergensis (Barendrecht, 1938)
  parvipalpis Edwards, 1925
  pusilla Edwards, 1925
  = sixi (Barendrecht, 1938) Note 7
  semiflava (Staeger, 1840)
                 
  Exechia Winnertz, 1863
  bicincta (Staeger, 1840)
  = interrupta (Zetterstedt, 1852)
  chandleri Caspers, 1987 N; will be formally added by Beuk & Chandler (in prep.)  Note 8
  cincta Winnertz, 1863
  confinis Winnertz, 1863
  contaminata Winnertz, 1863
  dizona Edwards, 1924
  dorsalis (Staeger, 1840)
  frigida (Boheman, 1865)
  fusca (Meigen, 1804)
  = lateralis (Meigen, 1818)
  = guttiventris (Meigen, 1830)
  lundstroemi Landrock, 1923
  = interrupta Landrock, 1909: preocc.
  nigrofusca Lundström, 1909
  nigroscutellata Landrock, 1912
  nitidicollis Lundström, 1913
  parva Lundström, 1909
  parvula (Zetterstedt, 1852) N; occurrence according to Fauna Europaea 
  pectinivalva Stackelberg, 1948 N; will be formally added by Beuk & Chandler (in prep.) 
  pseudofestiva Lackschewitz, 1937 N; occurrence according to Fauna Europaea 
  repanda Johannsen, 1912
  separata Lundström, 1912
  seriata (Meigen, 1830)
  = pallida (Stannius, 1831)
  spinigera Winnertz, 1863
  spinuligera Lundström, 1912
                 
  Exechiopsis Tuomikoski, 1966
  subg. Exechiopsis Tuomikoski, 1966
  aemula Plassmann, 1984 N; occurrence according to Fauna Europaea  Note 9
  fimbriata (Lundström, 1909)
  indecisa (Walker, 1856)
  = tenuicornis (Van der Wulp, 1859)
  intersecta (Meigen, 1818)
  pulchella (Winnertz, 1863)
  subulata (Winnertz, 1863)
  subg. Xenexechia Tuomikoski, 1966
  crucigera (Lundström, 1909)
  leptura (Meigen, 1830)
                 
  Pseudexechia Tuomikoski, 1966
  trivittata (Staeger, 1840)
                 
  Pseudobrachypeza Tuomikoski, 1966
  helvetica (Walker, 1856) N; will be formally added by Beuk & Chandler (in prep.) 
                 
  Rymosia Winnertz, 1863
  affinis Winnertz, 1863
  batava (Barendrecht, 1938)
  bifida Edwards, 1925
  = minor Landrock, 1925
  connexa Winnertz, 1863
  fasciata (Meigen, 1804)
  signatipes (Van der Wulp, 1859)
                 
  Stigmatomeria Tuomikoski, 1966
  crassicornis (Stannius, 1831)
  = obscura (Winnertz, 1863) Note 10
                 
  Synplasta Skuse, 1890
  = Gymnogonia Tuomikoski, 1966
  excogitata (Dziedzicki, 1910) N; will be formally added by Beuk & Chandler (in prep.) 
  gracilis (Winnertz, 1863)
                 
  Tarnania Tuomikoski, 1966
  fenestralis (Meigen, 1818)
  tarnanii (Dziedzicki, 1910)
 
  Mycetophilini
                 
  Dynatosoma Winnertz, 1863
  fuscicorne (Meigen, 1818)
  = praeusta (Meigen, 1830)
  reciprocum (Walker, 1848) N; will be formally added by Beuk & Chandler (in prep.) 
  thoracicum (Zetterstedt, 1838) N; will be formally added by Beuk & Chandler (in prep.) 
                 
  Epicypta Winnertz, 1863
  = Delopsis Skuse, 1890
  aterrima (Zetterstedt, 1852)
  limnophila Chandler, 1981 N; added by Beuk (1996h: 90, 202 Note 11
  scatophora (Perris, 1849)
                 
  Mycetophila Meigen, 1803
  = Fungivora Meigen, 1800: suppr.
  alea Laffoon, 1965
  = guttata Dziedzicki, 1884: preocc.
  attonsa Laffoon, 1957 N; occurrence according to Fauna Europaea 
  = uncta Plassmann, 1999
  autumnalis Lundström, 1909 N; occurrence according to Fauna Europaea 
  biusta Meigen, 1818
  blanda Winnertz, 1863
  britannica Laštovka & Kidd, 1975 N; occurrence according to Fauna Europaea 
  caudata Staeger, 1840 N; occurrence according to Fauna Europaea 
  cingulum Meigen, 1830
  = lunata Macquart, 1834
  confluens Dziedzicki, 1884
  confusa Dziedzicki, 1884 N; will be formally added by Beuk & Chandler (in prep.) 
  curviseta Lundström, 1911 N; will be formally added by Beuk & Chandler (in prep.) 
  dentata Lundström, 1915 N; occurrence according to Fauna Europaea 
  dziedzickii Chandler, 1977
  = obscura Dziedzicki, 1884: preocc.
  edwardsi Lundström, 1913
  = nebulosa Edwards, 1913: preocc.
  finlandica Edwards, 1913
  formosa Lundström, 1911
  fraterna Winnertz, 1863
  fungorum (De Geer, 1776)
  = punctata Meigen, 1804
  ichneumonea Say, 1823 N; will be formally added by Beuk & Chandler (in prep.) 
  idonea Laštovka, 1972 N; will be formally added by Beuk & Chandler (in prep.) 
  luctuosa Meigen, 1830
  marginata Winnertz, 1863
  mitis (Johannsen, 1912) N; added by Beuk (1996h: 90, 202
  morosa Winnertz, 1863 N; will be formally added by Beuk & Chandler (in prep.) 
  occultans Lundström, 1913 N; occurrence according to Fauna Europaea 
  ocellus Walker, 1848
  = dimidiata Staeger, 1840: preocc.
  = ocelus Walker, 1848: incorrect spelling
  perpallida Chandler, 1993 N; added by Beuk (1996h: 90, 202
  pictula Meigen, 1830
  = bimaculata (Fabricius, 1805): preocc.
  pumila Winnertz, 1863 N; will be formally added by Beuk & Chandler (in prep.) 
  rudis Winnertz, 1863 N; occurrence according to Fauna Europaea 
  ruficollis Meigen, 1818 N; will be formally added by Beuk & Chandler (in prep.) 
  sigillata Dziedzicki, 1884
  signata Meigen, 1830
  = lunata (Fabricius, 1805): preocc.
  signatoides Dziedzicki, 1884
  sordida Van der Wulp, 1874
  spectabilis Winnertz, 1863
  stolida Walker, 1856
  strigata Staeger, 1840
  strigatoides Landrock, 1927 N; occurrence according to Fauna Europaea 
  stylata (Dziedzicki, 1884)
  sumavica (Laštovka, 1963) N; will be formally added by Beuk & Chandler (in prep.) 
  trinotata Staeger, 1840
  unicolor Stannius, 1831
  uninotata Zetterstedt, 1852 N; occurrence according to Fauna Europaea 
  unipunctata Meigen, 1818 N; will be formally added by Beuk & Chandler (in prep.) 
  vittipes Zetterstedt, 1852
                 
  Phronia Winnertz, 1863
  = Telmatophilus Becker, 1908
  basalis Winnertz, 1863
  biarcuata (Becker, 1908)
  = praecox Edwards, 1925
  bicolor Dziedzicki, 1889
  = tarsata (Staeger, 1840): nomen dubium
  = crassipes Winnertz, 1863
  braueri Dziedzicki, 1889
  = annulata Winnertz, 1863: doubtful senior synonym
  = vittata Winnertz, 1863: doubtful senior synonym
  cinerascens Winnertz, 1863
  = truncata Winnertz, 1863
  conformis (Walker, 1856)
  coritanica Chandler, 1992 N; will be formally added by Beuk & Chandler (in prep.) 
  disgrega Dziedzicki, 1889 N; will be formally added by Beuk & Chandler (in prep.) 
  egregia Dziedzicki, 1889
  exigua (Zetterstedt, 1852)
  = rustica Winnertz, 1863
  flavipes Winnertz, 1863
  forcipata Winnertz, 1863
  forcipula Winnertz, 1863 Note 12
  gangnei Chandler, 1992 N; will be formally added by Beuk & Chandler (in prep.) 
  humeralis Winnertz, 1863
  = forcipula Winnertz, 1863: misident. sensu auct.
  mutabilis Dziedzicki, 1889
  nigricornis (Zetterstedt, 1852)
  = dubia Dziedzicki, 1889
  nitidiventris (Van der Wulp, 1859)
  = vitiosa Winnertz, 1863
  notata Dziedzicki, 1889 N; added by Beuk (2021b: 20
  obscura Dziedzicki, 1889
  obtusa Winnertz, 1863
  siebeckii Dziedzicki, 1889 Note 13
  signata Winnertz, 1863
  = austriaca Winnertz, 1863
  = flavicollis Winnertz, 1863
  strenua Winnertz, 1863
  tenuis Winnertz, 1863
                 
  Platurocypta Enderlein, 1910
  punctum (Stannius, 1831)
  testata (Edwards, 1925) N; will be formally added by Beuk & Chandler (in prep.) 
                 
  Sceptonia Winnertz, 1863
  concolor Winnertz, 1863
  costata (Van der Wulp, 1859)
  demeijerei Bechev, 1997 T; added by Bechev (1997: 180
  fumipes Edwards, 1925
  membranacea Edwards, 1925
  nigra (Meigen, 1804)
  = nitida (Meigen, 1830)
                 
  Trichonta Winnertz, 1863
  atricauda (Zetterstedt, 1852)
  = melanura (Staeger, 1840): misident sensu Van der Wulp & De Meijere, 1898
  fragilis Gagné, 1981 N; first listed by Beuk (2002e: 59
  melanura (Staeger, 1840)
  submaculata (Staeger, 1840) N; will be formally added by Beuk & Chandler (in prep.) 
  terminalis (Walker, 1856)
  vitta (Meigen, 1830)
  = submaculata (Staeger, 1840): misident. sensu Van der Wulp, 1874
                 
  Zygomyia Winnertz, 1863
  flaviventris Winnertz, 1863
  = nitida (Stannius, 1831): preocc.
  humeralis (Wiedemann, 1817)
  = canescens Winnertz, 1863
  notata (Stannius, 1831)
  pictipennis (Staeger, 1840)
  pseudohumeralis Caspers, 1980 N; occurrence according to Fauna Europaea  Note 14
  semifusca (Meigen, 1818) Note 15
  setosa Barendrecht, 1938
  valida Winnertz, 1863
  vara (Staeger, 1840)
 
MYCOMYINAE
                 
  Mycomya Rondani, 1856
  = Mycomyia Edwards, 1913: emend.
  subg. Calomycomya Väisänen, 1984
  pulchella (Dziedzicki, 1885)
  subg. Cymomya Väisänen, 1984
  circumdata (Staeger, 1840)
  = lucorum (Winnertz, 1863)
  subg. Mycomya Rondani, 1856
  annulata (Meigen, 1818)
  = incisurata (Zetterstedt, 1838)
  bicolor (Dziedzicki, 1885)
  cinerascens (Macquart, 1826)
  = alacris (Winnertz, 1863)
  flavicollis (Zetterstedt, 1852)
  frequens Johannsen, 1910 N; added by Beuk (1996h: 91, 202
  fuscata (Winnertz, 1863)
  insignis (Winnertz, 1863) N; will be formally added by Beuk & Chandler (in prep.) 
  marginata (Meigen, 1818)
  = limbata (Winnertz, 1863)
  neohyalinata Väisänen, 1984
  = hyalinata (Meigen, 1830): misident. sensu Zetterstedt, 1838
  nigricornis (Zetterstedt, 1852)
  nitida (Zetterstedt, 1852) N; will be formally added by Beuk & Chandler (in prep.) 
  occultans (Winnertz, 1863)
  ornata (Meigen, 1818)
  punctata (Meigen, 1804)
  tenuis (Walker, 1856)
  = apicalis (Winnertz, 1863)
  tumida (Winnertz, 1863) Note 16
  wankowiczii (Dziedzicki, 1885)
  = notabilis (Staeger, 1840): possible senior synonym
  = fissa (Lundström, 1911)
  winnertzi (Dziedzicki, 1885)
  subg. Mycomyopsis Väisänen, 1984
  affinis (Staeger, 1840)
  = flava (Winnertz, 1863)
  permixta Väisänen, 1984
  = trilineata (Zetterstedt, 1838): misident. sensu Dziedzicki, 1885
  subg. Neomycomya Väisänen, 1984
  fimbriata (Meigen, 1818)
                 
  Neoempheria Osten Sacken, 1878
  bimaculata (von Roser, 1840)
  pictipennis (Haliday, 1833) N; will be formally added by Beuk & Chandler (in prep.) 
  striata (Meigen, 1818)
 
SCIOPHILINAE
                 
  Acnemia Winnertz, 1863
  longipes Winnertz, 1863 N; added by Beuk (1996h: 91, 202
  nitidicollis (Meigen, 1818)
                 
  Allocotocera Mik, 1886
  = Eurycera Dziedzicki, 1885
  pulchella (Curtis, 1837)
  = silacea (Van der Wulp, 1887) Note 17
  = flava (Dziedzicki, 1889)
                 
  Anaclileia Meunier, 1904
  = Paraneurotelia Landrock, 1911
  dispar (Winnertz, 1863)
                 
  Azana Walker, 1856
  subg. Azana Walker, 1856
  anomala (Staeger, 1840)
                 
  Leptomorphus Curtis, 1831
  walkeri Curtis, 1831
                 
  Megalopelma Enderlein, 1910
  nigroclavatus (Strobl, 1909) N; added by Dek (2018: 85
                 
  Monoclona Mik, 1886
  = Staegeria Van der Wulp, 1876
  rufilatera (Walker, 1837)
  = halterata (Staeger, 1840)
  = forcipata Strobl, 1909: misident. sensu De Meijere, 1935
                 
  Neuratelia Rondani, 1856
  nemoralis (Meigen, 1818)
                 
  Phthinia Winnertz, 1863
  mira (Ostroverkhova, 1977)
  = humilis Winnertz, 1863: misident. sensu Hutson et al. 1980
                 
  Polylepta Winnertz, 1863
  guttiventris (Zetterstedt, 1852)
                 
  Sciophila Meigen, 1818
  = Lasiosoma Winnertz, 1863
  fenestella (Curtis, 1837)
  fractinervis Edwards, 1940 I; added by Chandler & Pijnakker (2009: 91
  hirta Meigen, 1818
  lutea Macquart, 1826
  nigronitida Landrock, 1925
  rufa Meigen, 1830
                 
  Speolepta Edwards, 1925
  leptogaster (Winnertz, 1863)
 
Doubtful species
 
  Mycetophila Meigen, 1803    
  flava Winnertz, 1863   Note 18
 
  Mycomya Rondani, 1856    
  subg. Mycomya Rondani, 1856  
  maculata (Meigen, 1804)   Note 19
 
  Phthinia Winnertz, 1863    
  humilis Winnertz, 1863   Note 20
 
  Sciophila Meigen, 1818    
  thoracica Staeger, 1840   Note 21
  = nigriventris Macquart, : misident.? 
 
Nomina dubia
  bicolor Macquart, 1834 (Mycetophila) Note 22
  flaviventris Van der Wulp, 1859 (Mycetophila) Note 22
  lineola Meigen, 1818 (Mycetophila) Note 23
  ruficauda Van der Wulp, 1874 (Brachycampta) Note 22
  rufithorax Van der Wulp, 1874 (Exechia) Note 24
  schummeli Stannius, 1831 (Mycetophila) Note 25
 
Deleted species
 
  Mycetophila Meigen, 1803  
  gratiosa Winnertz, 1863   Note 26
  lunata Meigen, 1804   Note 27
  xanthopyga Winnertz, 1863   Note 26
 
  Mycomya Rondani, 1856  
  subg. Mycomya Rondani, 1856  
  levis (Dziedzicki, 1885)   Note 28
 
Notes
Note 1 Based on the illustrations in Barendrecht (1938), Zaitzev et al. (2005) suspect that Barendrecht’s record of dispecta actually refers to gusakovae Zaitzev, 1994.
Note 2 Synonymy following Fauna Europaea.
Note 3 Erroneously listed as new for the Netherlands by W. van Steenis & Jacobusse (2017: 10).
Note 4 Material examined: LIMBURG: Meinweg (3.2 km NE of Herkenbosch: along Bosbeek near post 405, GPS: 51.1706N 6.0969E: 2 [m], 30.ix-10.x.2016, leg. NHM Maastricht MT [NHMM alc]).
Note 5 This is the species in the female sex sensu Plassmann (1970: 397).
Note 6 First recorded by De Meijere (1935: 194) but later deleted (De Meijere 1939a: 123) because the material supposedly belonged to other species (unspecified) or because it could not be identified. At present it is considered to be the probable senior synonym of bergensis (see Fauna Europaea).
Note 7 This possible synonymy needs to be confirmed.
Note 8 Material examined for original record: NOORD-BRABANT: Udenhout (de Brand, 1 [m]: 20-27.x.1990, leg. Ins.W.G. KNNV-Tilburg MT [BEUK]). Additional material examined: LIMBURG: Meinweg (3.2 km NE of Herkenbosch: along Bosbeek near post 405 [GPS 51.1706N 6.0969E]: 5 [m]: 1-10.xi.2015, leg. P. Thomas, Malaise trap [NHMM ALC]).
Note 9 Material examined: LIMBURG: Meinweg (3.2 km NE of Herkenbosch: along Bosbeek near post 405, GPS: 51.1706N 6.0969E: 1 [m], 20-30.vii.2016, leg. NHM Maastricht MT [NHMM alc]).
Note 10 Synonymy following Fauna Europaea.
Note 11 Old Dutch material recorded as E. scathophora may in fact belong to this species, e.g., E. scathophora Perris sensu Edwards is E. limnophila Chandler.
Note 12 So far all material recorded under this name that could be checked belonged to P. humeralis. It may wel be that this species does not occur in the Netherlands.
Note 13 Material examined: GELDERLAND: Wageningen (Nassauweg [GPS: 51.966N, 5.673E], under light dome: 1 [m], 15.ii.2022, leg. D. Belgers [NHMM ALC]).
Note 14 ZUID-HOLLAND: Wassenaar (Meijendel, duin [RD 85.7-463.3]: 1 [m], 30.viii-5.ix.2000, leg. H.J. Prijs MT [NHMM ALC]).
Note 15 This species had already been transferred from Mycetophila to Zygomyia by Zaitzev (1989a).
Note 16 Some earlier published records of this species (Väisänen, 1982b: 41; 1984: 215; Hackman et al., 1988: 229) were based on an old record by Van der Wulp (1877: 168; material later referred to M. (M.) ornata) and on doubtfully identified females (Väisänen; 1984: 57). However, recently collected material of this species was examined so it is herewith confirmed for the list.
Note 17 This species was synomynised by Edwards (1925: 557) and this synonymy was followed by De Meijere (1939b). However, since Hackman et al. (1988) listed it as valid species it was included in by Beuk (2002e).
Note 18 Doubtfully recorded from the Netherlands by Hackman et al. (1988: 269) and followed by Fauna Europaea. No other published records are known.
Note 19 De Meijere (1939a: 123) deleted the species from the Dutch list but a number of recent publications (Väisänen, 1984: 212; Hackman et al., 1988: 226) still list it as Dutch. No examined material is mentioned in any these publications so the record is considered doubtful.
Note 20 The original record of this species by Schmitz (1929) could not be verified, other material belonged to mira.
Note 21 Listed under the name S. nigriventris with S. thoracica as synonym up to Van der Wulp & De Meijere (1898: 11), but not mentioned since.
Note 22 This is an unrecognized species in the Allodia genus group according to Hackman et al. (1988).
Note 23 This is an unrecognized species of the Mycetophila ruficollis group and Van der Wulp (1877: 151) treated it as senior synonym of M. ruficollis. It was still treated as valid species by Van der Wulp & De Meijere (1898: 9), Barendrecht (1938b: 52) and De Meijere (1939b: 143) but now it is considered as nomen dubium by Hackman et al. (1988: 279).
Note 24 This is an unrecognized species in the Exechia genus group according to Hackman et al. (1988).
Note 25 This is an unrecognized species provisionally placed in the Exechia genus group but does not seem to belong there according to Hackman et al. (1988).
Note 26 First recorded by Van der Wulp (1874: 120) and records repeated by Van der Wulp (1877) and Van der Wulp & De Meijere (1898), but later deleted by De Meijere (1939a: 123) as belonging to unspecified other species.
Note 27 First recorded by Van der Wulp (1877: 155) and records repeated by Van der Wulp & De Meijere (1898: 9), but later deleted by De Meijere (1939a: 123) as belonging to unspecified other species.
Note 28 First recorded by De Meijere (1935: 194) but later deleted (De Meijere 1939a: 123) because the material supposedly belonged to unspecified other species or because it could not be identified.
 
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Allodia was the last (sub)genus to be edited on 15-11-2023 15:30
Phronia siebeckii was the last (sub)species to be edited on 13-05-2022 15:09















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